"Bart@nerdster.org"
is Bart.
We see "Bart@nerdster.org" because
Bart used the Nerdster to sign and publish the follow statement.
(Same dude as "Bart@ebay.com" or
"Bart@discord.com" would be.)
"Milhouse" is Milhouse (simply, with no @domain) because that's who Bart wants to follow: a person, not an account.